Applied and Environmental Microbiology

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Editor in Chief L. Nicholas Ornston (2011) Yale University Minireview Editor Daniel J. Arp (2009) Oregon State University Editors Axel A. Brakhage (2011) HKI and University of Jena Daniel Cullen (2012) USDA Forest Products Laboratory Harold L. Drake (2012) University of Bayreuth Katharine J. Gibson (2011) DuPont Experiment Station Lone Gram (2011) Danish Institute for Fisheries Research Mansel W. Griffiths (2010) University of Guelph Jo Handelsman (2010) University of Wisconsin Sophia Kathariou (2012) North Carolina State University Michael J. Larkin (2012) The Queen's University of Belfast Laura G. Leff (2011) Kent State University Frank E. Löffler (2013) Georgia Institute of Technology Charles R. Lovell (2010) University of South Carolina Ellen L. Neidle (2012) University of Georgia Douglas H. Ohlendorf (2011) University of Minnesota Matthew R. Parsek, Cover Editor (2009) University of Washington Donald W. Schaffner (2010) Rutgers, The State University of New Jersey Alfred M. Spormann (2011) Stanford University Marylynn V. Yates (2009) University of California – Riverside Jizhong Zhou (2013) University of Oklahoma Chairman, Publications Board Thomas E. Shenk Director, Journals Barbara M. Goldman Production Editor Barbara S. Slinker Assistant Production Editor Ellie Ghatineh

Editorial Board Yousef Abu Kwaik (2008) Luis A. Actis (2008) Eric Altermann (2009) Gary Andersen (2010) Thomas Andlid (2010) Adam Arkin (2009) Beate Averhoff (2010) Andrea Azcarate (2009) Felix J. Baerlocher (2009) Michael Bagdasarian (2009) Bert Bago (2010) Christopher E. Bagwell (2008) Alan Bakalinsky (2009) Katherine H. Baker (2009) Jo'zsef Baranyi (2010) Tamar Barkay (2008) Douglas H. Bartlett (2008) Florian Bauer (2009) Gwyn A. Beattie (2008) Harry Beller (2009) Tahia Benitez (2008) Andrew K. Benson (2009) Peter L. Bergquist (2009) Colin Berry (2008) Tom Besser (2010) Arvind A. Bhagwat (2009) K. Johanna Bjorkroth (2010) Rafael Blasco (2010) David E. Block (2010) Paul L. E. Bodelier (2008) Antje Boetius (2008) Eckhard Boles (2009) Paola Bonfante (2009) Violaine Bonnefoy (2010) Kathryn J. Boor (2010) Katherine A. Borkovich (2009) James Borneman (2008) Dulal Borthakur (2009) Thomas L. Bott (2009) Kostas Bourtzis (2009) Gesche Braker (2008) Michael G. Bramucci (2008) Byron Brehm-Stecher (2010) Frederick Breidt, Jr. (2010) Neil C. Bruce (2010) George Bullerjahn (2010) William Burkhardt III (2010) H. J. Busscher (2010) Mark P. Buttner (2009) Murulee Byappanahalli (2009) Douglas R. Call (2010) Terri A. Camesano (2009) Russell W. Carlson (2010) Michael J. Carter (2010) Jinru Chen (2008) Wilfred Chen (2009) Michael L. Chikindas (2010) John D. Coates (2008) Luca Cocolin (2010) James Cole (2010) Mary Lynne Perille Collins (2008) Michael T. Collins (2010) James M. Cregg (2009) Patricia Cruz (2010) Tom Curtis (2010) Lori L. Daane (2008) Rolf Daniel (2009) Steven L. Daniel (2008) Michelle Danyluk (2010) Atin R. Datta (2010) Frank B. Dazzo (2010) Angelo DePaola, Jr. (2009) Anne Desjardins (2008) Ronald P. de Vries (2008) Thomas J. DiChristina (2008) Alan DiSpirito (2008) Jan Dolfing (2010) Daniel P. Dougherty (2009) Irina S. Druzhinina (2008) Nicole Dubilier (2010) Siobain Duffy (2010) Peter Dürre (2008) Nancy DuTeau (2008) Richard Eaton (2010) Leo Eberl (2008) T. S. Edrington (2010) Henry L. Ehrlich (2010) Christopher Elkins (2008) David Emerson (2009) Scott A. Ensign (2008) K.-D. Entian (2010) Slava Epstein (2010) Danilo Ercolini (2009) David Faguy (2008) Joseph O. Falkinham III (2010) Brian Federici (2009) Juan Ferré (2009) Matthew W. Fields (2008) Turlough Finan (2010) Steven E. Finkel (2010) Madilyn Fletcher (2008) Christopher Francis (2010) James A. Fraser (2009) Herbert L. Fredrickson (2010) Masao Fukuda (2009) Clay Fuqua (2008) Manuela Giovannetti (2009) Heidi Goodrich-Blair (2009) Lawrence Goodridge (2009) Joerg Graf (2008) Irene R. Grant (2010) Jean Guard-Bouldin (2010) Mary Lou Guerinot (2008) Martin W. Hahn (2010) Ken Hammel (2010) Theo A. Hansen (2010) Hauke Harms (2010) Steven Harris (2008) Valerie J. (Jody) Harwood (2009) Terry C. Hazen (2010) Qiang He (2010) Zhili He (2010) Hermann Heipieper (2008) Alfredo Herrera-Estrella (2008) Russell T. Hill (2008) Dallas G. Hoover (2009) David J. Hopper (2010) Hor-Gil Hur (2009) Fumio Inagaki (2010) Douglas Inglis (2009) Karl-Erich Jaeger (2008) Janet K. Jansson (2010) Lee-Ann Jaykus (2009) Tom Jeffries (2008) Paul R. Jensen (2008) Peter Jensen (2008) Carlos A. Jerez (2008) Mike Jetten (2010) D. Barrie Johnson (2008) Yves Jouanneau (2009) Ari Jumpponen (2009) Juan Luis Jurat-Fuentes (2009) David Kaplan (2010) Levente Karaffa (2009) Charles W. Kaspar (2010) Jay D. Keasling (2008) James E. Keen (2010) Martin Keller (2009) John J. Kelly (2009) Robert M. Kelly (2008) Angela Kent (2010) Lee Kerkhof (2009) Philip J. Kersten (2010) David L. Kirchman (2008) Mary Jo Kirisits (2010) Maia Kivisaar (2010) Staffan Kjelleberg (2008) Martin G. Klotz (2008) Allan Konopka (2008) Joel E. Kostka (2010) George Kowalchuk (2009) Lee Krumholz (2009) Indira Kudva (2010) Kirsten Kuesel (2010) Timothy M. LaPara (2009) Laurie LaPat-Polasko (2009) Gisèle LaPointe (2010) Jared Leadbetter (2010)

Yin-Won Lee (2010) Mary Beth Leigh (2010) Jeffrey T. LeJeune (2009) Michael Lemke (2008) Celeste Linde (2008) Markus Linder (2008) Nic D. Lindley (2010) John T. Lisle (2009) Wen-Tso Liu (2010) Jon Lloyd (2010) G. T. Macfarlane (2008) Robert Mach (2009) Eugene L. Madsen (2008) Walter F. Mahaffee (2008) Mike Manefield (2008) Helene Marquis (2009) Robert E. Marquis (2008) Terence L. Marsh (2008) Andrew P. Martin (2008) Maria Esperanza Martínez-Romero (2010) Jan Martinussen (2008) Juergen Marxsen (2008) Luke Masson (2008) Harold D. May (2010) Alejandro Mazzotta (2008) J. Vaun McArthur (2008) Timothy R. McDermott (2008) Katherine McMahon (2009) Brian McSpadden-Gardner (2010) Rick Meinersmann (2009) Aubrey F. Mendonca (2009) Jianghong Meng (2009) Eric S. Miller (2008) Robert V. Miller (2008) Patricia D. Millner (2009) Donald K. Milton (2009) Kiwamu Minamisawa (2008) William W. Mohn (2008) Sylvain Moineau (2008) Istvan Molnar (2008) Melanie Mormile (2010) Volker Müller (2010) Peter Muriana (2008) Yuji Nagata (2009) István Nagy (2010) Cindy H. Nakatsu (2010) Diane G. Newell (2010) Walter G. Niehaus, Jr. (2010) Brendan A. Niemira (2009) Hideaki Nojiri (2009) Kenneth M. Noll (2008) Norbert Nowotny (2009) George-John Nychas (2009) Mark R. O'Brian (2009) Kevin P. O'Connell (2008) Ronald S. Oremland (2009) Mark Osborn (2010) Joerg Overmann (2010) Rebecca Ehrlich Parales (2008) Matthew A. Parker (2008) John H. Paul (2008) Jakob Pernthaler (2009) M. Julia Pettinari (2008) Flynn Picardal (2010) Roger W. Pickup (2010) James L. Pinckney (2008) Anton F. Post (2010) Rolf Prade (2009) Jim I. Prosser (2008) Cheryl L. Quinn (2009) John P. Quinn (2009) Jos M. Raaijmakers (2008) Fredrick A. Rainey (2008) Gerald R. Reeck (2010) Bernd Rehm (2009) Gregor Reid (2009) Walter Reineke (2010) Geoffrey D. Robson (2008) Neil Rowan (2008) Edward G. Ruby (2010) Michael J. Sadowsky (2010) Badal C. Saha (2010) Andrew D. Sails (2010) Sima Sariaslani (2009) Brian Sauders (2010) Patrick D. Schloss (2009) Andreas Schmid (2008) Janet Schottel (2008) Andreas Schramm (2010) Kate M. Scow (2008) Bernhard Seiboth (2008) Verena Seidl (2009) Gordon Shephard (2010) Wenyuan Shi (2008) Simon D. Silver (2008) Anne Simon (2008) Pascal Simonet (2009) Randall S. Singer (2009) Brajesh Singh (2010) Kaarina Sivonen (2008) Joan L. Slonczewski (2008) Kornelia Smalla (2009) Hauke Smidt (2008) Patricia A. Sobecky (2010) John Spear (2010) Andrew Spiers (2010) Dirk Springael (2009) Ewald Srebotnik (2008) Eric Stabb (2010) Lucas J. Stal (2009) Alfons J. M. Stams (2010) Robert J. Steffan (2010) Craig Stephens (2009) John Stolz (2010) Wolfgang Streit (2009) Herbert Strobel (2008) Joseph Sturino (2010) Marcelino Suzuki (2010) Ken Takai (2010) Gerald W. Tannock (2008) Ron Teather (2010) Tuula Teeri (2008) Andreas Teske (2009) Chris M. Thomas (2009) Michael G. Thomas (2009) Ian P. Thompson (2008) Tim Tolker-Nielsen (2009) Eva M. Top (2009) Effie Tsakalidou (2010) Masataka Tsuda (2009) Jan van Beilen (2008) Jos Vanderleyden (2010) Alain Vertes (2010) Anne K. Vidaver (2008) Gerrit Voordouw (2008) Adrián A. Vojnov (2010) Carmen Wacher (2010) Michael Wagner (2010) Irene Wagner-Döbler (2008) Suzanne Walker (2009) Judy D. Wall (2009) Peng G. Wang (2009) Todd Ward (2010) Kazuya Watanabe (2008) Bart Weimer (2010) Ronald Weiner (2008) Elizabeth M. Wellington (2010) Alan Welman (2009) Andrew S. Whiteley (2008) Marvin Whiteley (2010) Giovanni Widmer (2008) Martin Wiedmann (2008) Steve Wilhelm (2009) Joanne Willey (2008) Rolf-M. Wittich (2008) Mark J. Wolcott (2010) K. Eric Wommack (2009) Thomas K. Wood (2008) Randy Worobo (2010) Michael Wyman (2009) Dong Xu (2008) Jian Xu (2010) Luying N. Xun (2008) Chuanlun Zhang (2009) Erwin G. Zoetendal (2009) Angeles Zorreguieta (2010)

Applied and Environmental Microbiology, a publication of the American Society for Microbiology (ASM), 1752 N St., N.W., Washington, DC 20036-2904, is devoted to the advancement and dissemination of applied knowledge as well as ecological knowledge, both applied and fundamental, concerning microorganisms.